What the work claims
Raup and Sepkoski claimed that the dinosaur extinction 66 mya was part of a recurring cycle of mass extinctions occurring every 26 million years.
How it was done
Raup and Sepkoski used statistical analysis of the fossil record to detect periodicity in extinction events.
Raup was born on 24 April 1933 and raised in Boston, and he came late to the fossil record, having had very little contact with it as a child; he preferred skiing and camping. His first mentor was John Clark, a vertebrate paleontologist and sedimentologist at the University of Chicago. He began at Colby College in Maine, transferred to Chicago for his Bachelor of Science, and took his MA and PhD at Harvard, majoring in geology with a focus on paleontology and biology. He taught at Caltech, Johns Hopkins and the University of Rochester, and was a curator and Dean of Science at the Field Museum of Natural History in Chicago. He retired in 1994 to Washington Island in northern Lake Michigan and died on 9 July 2015. The asteroid 9165 Raup is named for him, and he was elected to the American Academy of Arts and Sciences in 1996 and the American Philosophical Society in 2002.
What holds up
The statistical detection of periodicity in extinction intensity across the Phanerozoic fossil record remains a documented analytical result from their 1984 PNAS paper.
The 1984 paper is titled “Periodicity of extinctions in the geologic past” and appeared in Proceedings of the National Academy of Sciences. It followed their 1982 paper in Science, “Mass extinctions in the marine fossil record”. Raup had also published “Size of the Permo-Triassic Bottleneck and Its Evolutionary Implications” in Science in 1979.
What does not
The 26-million-year cycle has not been confirmed as causal or physical. The work does not identify a mechanism, nor does it rule out statistical artefact or sampling bias.
Why it matters beyond the lab
It forced geology and evolutionary biology to confront whether extinction is driven by internal biosphere dynamics or external astrophysical triggers — a question still unresolved.
Raup returned to the subject in book form. The Nemesis Affair: A Story of the Death of Dinosaurs and the Ways of Science appeared in 1986 and was revised and expanded in 1999. Extinction: Bad Genes or Bad Luck? followed in 1992, and in 1994 he published “The Role of Extinction in Evolution” in PNAS.
Raup was not only an extinction specialist. He studied the fossil record and the diversity of life on Earth, and his earliest listed papers are on the form of shells: “Computer as aid in describing form in gastropod shells” in Science in 1962 and “Geometric analysis of shell coiling: general problems” in the Journal of Paleontology in 1966. Late in life he helped the Santa Fe Institute develop methods for the evolutionary exploration of morphospace.
Is it worth your time
Yes — it redefined how palaeontologists interpret deep-time patterns, but only as a hypothesis grounded in one dataset and method.





